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This study describes the comparative evaluation under monotonic and cycled load (butt and overlap welded) of dissimilar steels; structural ASTM A537 (I) and stainless ASTM A240 (304L) through GMAW, Argon as protecting gas and ASTM A240 (308L) as a supplier material. Microstructures were contrasted in different zones of each joint, focus on the Heat Affected Zone (HAZ) and fusion lines. The following mechanical tests were compared between both welding joint (WJ): Vickers hardness profile, tension, bending, and axial fatigue. A statistically based model was determined for each configuration that allows predicting the fatigue life of Wöhler field and also the fatigue factor. Vickers profile show high values of microhardness in the HAZ, near the fusion line between weld and stainless. Tension and axial fatigue tests indicated similar behavior between WJ and structural steel (butt joint); and similar behavior between WJ and stainless (overlap joint). Dissimilar unions (butt and overlap) have mechanical and microstructure properties under monotonic and cyclic loading, which can be considered adequate to withstand the mechanical requirements in service conditions, despite relatively high values of hardness in the HAZ.